#NEXUS [MacClade 4.06 registered to torsten dikow, fmnh] BEGIN DATA; DIMENSIONS NTAX=175 NCHAR=220; FORMAT SYMBOLS= " 0 1 2 3 4 5 6 7 8" MISSING=? GAP=- ; CHARSTATELABELS 1 shape_of_head / wider_than_high 'circular_(as_wide_as_high)', 2 compound_eye_separation / 'females_+_males_dichoptic' 'females_dichoptic,_males_holoptic', 3 'vertex shape between compound eyes (in dichoptic females)' / vertex_and_compound_eyes_at_same_level 'vertex_slightly_depressed_(less_than_60¡_angle_on_median_margin_of_compound_eye)' 'vertex_sharply_depressed_(90¡_angle_on_lateral_margin_of_compound_eyes)', 4 extent_of_facial_swelling / not_developed lower_facial_margin_slightly_developed extending_over_lower_facial_half extending_over_lower_three_quarters extending_over_entire_face, 5 division_of_face / 'entire,_no_division_evident' 'divided,_distinct_ventral_and_dorsal_half_developed' 'divided,_prominent_ventral_part_and_narrow_dorsal_part_developed', 6 mystax / absent present, 7 extent_of_mystax / restricted_to_lower_facial_margin extending_over_lower_half extending_over_lower_three_quarters extending_over_entire_face, 8 shape_of_mystax / setae_loosely_arranged 'setae_tightly_packed,_forming_a_distinct_shield', 9 clypeus / 'inverted_U-shaped_sclerite' 'inverted_U-shaped_sclerite,_dorsal_half_slcerotised_to_form_a_plate' 'sclerites_extended_medially,_nearly_forming_a_single_plate', 10 'clypeal-facial margin' / forming_a_distinct_ridge 'smooth,_not_particularly_pronounced', 11 maxillary_palpus / 'one-segmented' 'two-segmented', 12 sensory_pit_in_distal_palpomere / absent present, 13 distal_palpomere / cylindrical 'laterally_compressed_(leave-like)', 14 stipites_fusion / fused_entirely_medially 'fused_medially,_but_with_V-shaped_indentation' divided_medially not_slcerotised, 15 stipites_fusion_to_postgenae / separated fused, 16 postgenae_medial_projection / absent present, 17 shape_of_proboscis / straight angled_evenly_ventrally angled_sharply_ventrally distal_tip_angled_evenly_dorsally, 18 'cross-section of prementum proximally' / circular 'dorso-ventrally_flattened' laterally_compressed square not_sclerotised, 19 postmentum_ventral_development / ventral_margin_entirely_smooth median_groove_only_in_distal_part median_groove_on_entire_length, 20 postmentum_ventral_margin / 'plate-like,_straight' 'plate-like,_arched', 21 'post- and prementum fusion' / not_fused entirely_fused, 22 dorsal_flange_medially_on_prementum / absent present, 23 labellae / 'well-developed_and_separated_from_prementum' 'reduced,_fused_to_prementum_only_ventrally' 'reduced,_fused_to_prementum_entirely', 24 apex_of_labellae / rounded pointed blunt 'enlarged,_pseudotracheae_present', 25 length_of_labella / 'small,_only_forming_distal_tip_of_proboscis' 'large,_occupying_from_one-third_to_one-half_the_length_of_proboscis', 26 extent_of_sensory_setae_on_prementum_and_labella / setae_on_prementum_and_labella setae_restricted_to_labella, 27 maxillae_length / shorter_than_labrum longer_than_labrum, 28 shape_of_maxillae / same_height_throughout 'proximally_high,_distal_tip_distinctly_narrower' tapering_gradually_towards_tip, 29 tip_of_maxillae / blunt pointed, 30 hypopharynx / only_lightly_sclerotised heavily_sclerotised, 31 hypopharynx_length / 'short,_much_shorter_than_labium' 'long,_as_long_as_labium', 32 shape_of_sclerotised_hypopharynx / 'parallel-sided_throughout,_only_distal_tip_suddenly_pointed' 'parallel-sided_in_proximal_half,_distal_half_much_narrower' tapering_gradually_towards_distal_tip, 33 hypopharynx_dorsal_protuberances / absent 'present,_seta-like_spicules_spaced_close_together' 'present,_seta-like_spicules_spaced_far_apart', 34 length_of_labrum / 'long,_as_long_as_labium_(or_nearly_so)' 'short,_at_most_half_the_length_of_labium', 35 'dorso-posterior margin of cibarium' / 'simple,_no_particular_ridge_connecting_cornua_developed' 'one_transverse_ridge_connecting_cornua,_cornua_originating_from_ridge' 'one_transverse_ridge_connecting_cornua,_cornua_not_originating_from_ridge', 36 shape_of_cibarium / trapezoidal smoothly_rounded dorsally_narrow_and_ventrally_broad, 37 development_of_cornua_of_cibarium / 'well-developed_in_antero-posterior_orientation' 'well-developed_in_dorso-ventral_orientation' reduced_or_absent, 38 median_longitudinal_ridge_on_cibarium / absent present, 39 pharyngeal_pump / absent present, 40 pharyngeal_pump_development / laterally_without_sclerotised_ring 'laterally_with_sclerotised_ring,_but_medio-anteriorly_unsclerotised' with_entire_sclerotised_ring, 41 'pharyngeal pump shape (dorsal view)' / circular triangular, 42 dorsal_margin_of_postocciput / 'simple,_no_projections' triangular_projection_present, 43 shape_of_occipital_foramen / a_single_foramen two_distinct_foramina, 44 anterior_tentorial_pits / 'small,_slit-like,_inconspicuous,_ventrally' 'well-developed,_conspicuous,_antero-ventrally' 'well-developed,_conspicuous,_antero-medially' absent, 45 facial_setae / only_circular_setae 'circular_and_dorso-ventrally_flattened_setae_laterally_in_males_and_females' 'circular_and_dorso-ventrally_flattened_mystacal_setae_in_males' 'circular_and_dorso-ventrally_flattened_mystacal_setae_in_males_and_females', 46 size_of_anterior_ommatidiae / all_same_size at_least_some_median_ommatidiae_distinctly_larger, 47 postgenae / 'posterior_margin_simple,_smooth' 'posterior_margin_with_large_flange-like_projection', 48 'shape of frons (at level of antennal insertion)' / 'more_or_less_parallel-sided' markedly_and_suddenly_diverging_laterally markedly_approximating_medially, 49 'macrosetae on lateral margin of frons (dorsal to antennal insertion)' / absent present, 50 'elevated ridge medially on frons (dorsal to antennal insertion)' / absent 'present,_connecting_antennae_and_ocellar_triangle', 51 position_of_antennae_on_head / in_ventral_half in_dorsal_half, 52 distance_between_scapes / 'scapes_close_together,_touching' scapes_separated, 53 elevation_of_antennal_insertion / not_elevated 'elevated_above_eye_margin_(in_lateral_view)_with_distinct_protuberance', 54 shape_of_postpedicel / tapering_distally 'cylindrical,_same_diameter_throughout' 'medially_broadest,_long' 'elongated_and_distinctly_clubbed_distally_(Mydidae)' proximal_bulb_with_elongated_distal_part 'medially_broadest,_short', 55 setae_on_dorsal_postpedicel / absent present, 56 sclerotisation_of_postpedicel / entirely_sclerotised cylindrical_proximal_part_and_bulbous_distal_part_separated_by_unsclerotised_area, 57 'apical ''seta-like'' sensory element' / apically_on_stylus apically_in_cavity_on_stylus 'sub-apically_in_cavity_on_postpedicel' 'sub-apically_laterally_on_stylus' apically_in_cavity_on_postpedicel 'sub-apically_laterally_on_postpedicel', 58 stylus / three_elements two_elements one_element 'reduced,_only_apical_''seta-like''_sensory_element_present', 59 setae_on_stylus / absent present_on_anterior_surface_only present_on_all_surfaces, 60 ocelli_on_ocellar_triangle / all_three_ocelli_on_a_single_ocellar_triangle 'anterior_ocellus_separate,_situated_anteriorly', 61 development_of_posterior_ocelli / circular_or_slightly_oval distinctly_elongated, 62 'ocellar (oc) setae' / absent setae_only macrosetae_and_setae, 63 shape_of_postcranium / flat_to_convex concave, 64 'shape of posterior compound eye margin (lateral view)' / straight_or_lightly_curved_anteriorly_throughout sinuate_in_ventral_quarter distinctly_sinuate_in_ventral_half, 65 'macrosetae on median occipital sclerite (m ocp scl)' / absent two_distinct_setae more_than_two_setae, 66 'postocular (pocl) setae' / absent setae_only macrosetae_and_setae, 67 'shape of postocular (pocl) setae' / straight_or_only_lightly_angled_anteriorly angled_distinctly_anteriorly_in_distal_half, 68 division_of_pronotum / only_a_single_sclerite_present 'divided_into_ante-_and_postpronotum', 69 antepronotum_setae / absent setae_only macrosetae_and_setae, 70 'lateral postpronotal (pprn) setae' / absent setae_only macrosetae_and_setae, 71 dorsal_fusion_of_postpronotal_lobes_with_scutum / entirely_fused 'only_partly_fused,_lobes_elevated_above_scutum_posteriorly', 72 postpronotal_lobes / confined_to_lateral_margin_of_scutum 'extending_medially,_but_not_anteriorly' 'extending_medially_and_anteriorly,_nearly_touching_medially', 73 'dorsal postpronotal (pprn) setae' / absent setae_only macrosetae_and_setae, 74 lateral_fusion_of_prosternum_to_proepisternum / 'fused,_broad_prosternum' 'fused,_prosternum_narrow_above_pro_coxa' 'separated,_sclerite_triangular,_pointed_dorsally' 'separated,_sclerite_square_to_rectangular,_straight_dorsally' 'separated,_prosternum_square_and_proepisternum_projecting_medially_(nearly_touching_prosternum)', 75 dorsal_margin_of_prosternum / not_elevated 'distinct_flange-like_projection', 76 shape_of_propresternum / 'arrow-like_sclerite' square_sclerite triangular_sclerite, 77 development_of_cervical_sclerite / not_extending_medially_to_cover_sensory_area_on_propresternum extending_medially_to_cover_sensory_area_on_propresternum_partly, 78 anterior_anepisternal_setae / absent erect_setae 'erect_to_appressed,_anteriorly_directing_setae', 79 anterior_proepimeral_setae / absent erect_setae 'erect_to_appressed,_anteriorly_directing_setae', 80 'macrosetae on supero-posterior anepisternum' / absent present, 81 setae_on_posterior_basalare / absent present, 82 anatergal_setae / absent setae_only 'short,_stout_macrosetae', 83 katatergal_setae / absent setae_only macrosetae_and_setae, 84 katatergite_shape / more_or_less_flat_sclerite 'cone-shaped_sclerite', 85 'setae on posterior meron + metanepisternum' / absent setae_only at_least_one_macroseta_among_setae, 86 'setae on supero-posterior anepimeron' / absent present at_least_one_macroseta_among_setae, 87 'short, stout, erect macrosetae on anterior mesonotum' / absent two_macrosetae_present, 88 'notopleural (npl) setae' / absent one_seta two_setae three_or_more_setae 'single,_short_stout_macroseta', 89 'transverse (trn) suture' / 'entirely_absent_(not_discernible)' at_least_on_lateral_scutum_distinct, 90 'supra-alar (spa) setae' / absent present Hardy_1930_Dikow, 91 'postalar (pal) setae' / absent present, 92 'presutural dorsocentral (dc) setae' / absent present, 93 'presutural acrostichal (acr) setae' / absent present, 94 'postsutural dorsocentral (dc) setae' / absent present, 95 base_of_scutal_setae / setae_without_large_sockets setae_with_large_sockets, 96 'development of medial setae on posterior scutum (between dc setae)' / absent 'present,_directed_posteriorly' 'present,_directed_anteriorly' 'present,_appressed_to_surface,_directed_posteriorly', 97 size_of_scutellum / 'small,_mesopostnotum_visible_in_dorsal_view' 'large,_mesopostnotum_not_visible_in_dorsal_view', 98 'apical scutellar (ap sctl) setae' / absent setae_only macrosetae_and_setae, 99 'discal scutellar (ds sctl) setae' / absent present, 100 'coxae and metkatepisternum (in ventral view)' / metkatepisternum_small_and_not_visible_between_mes_and_met_coxae metkatepisternum_large_and_visible_between_mes_and_met_coxae, 101 metkatepisternum_setae / absent present, 102 postmetacoxal_bridge / 'absent,_postmetacoxal_area_entirely_membranous' 'partly_present_laterally,_membranous_area_medially' 'present,_visible_suture_between_lateral_halves' 'present,_entirely_sclerotised', 103 shape_of_setae_on_legs / only_circular_setae_present 'dorso-ventrally_flattened_setae_and_circular_setae_present', 104 lateral_depression_on_prothoracic_coxa / absent present, 105 'setae on antero-ventral prothoracic tibia' / absent at_least_three_setae_present, 106 projections_on_prothoracic_tibia / absent 'small_S-shaped_movable_spur,_originating_postero-ventrally' 'large_spine,_originating_antero-ventrally', 107 'base of spine/spur on prothoracic tibiae' / spur_originating_directly_from_tibiae spur_originating_from_projection_on_tibiae, 108 proximal_prothoracic_tarsomere / cylindrical cylindrical_with_short_spikes_proximally 'expanded_proximo-ventrally_with_stout_spikes', 109 'orientation of pro- and mesothoracic coxae' / distinctly_directed_posteriorly 'more_or_less_directed_ventrally_(sometimes_even_anteriorly)', 110 'row of macrosetae on antero-dorsal mesothoracic tibia' / absent one_seta_present several_setae_present, 111 orientation_of_metathoracic_coxae / distinctly_directed_ventrally_to_posteriorly directed_anteriorly, 112 setae_on_lateral_metathoracic_coxae / absent setae_only macrosetae_and_setae, 113 protuberance_on_anterior_metathoracic_coxae / absent 'present,_protuberance_blunt_distally' 'present,_peg-like_protuberance_pointed_distally', 114 setae_on_median_metathoracic_trochanter / absent setae_only macrosetae_and_setae, 115 shape_of_median_metathoracic_trochanter / 'cylindrical,_medially_without_protuberances' medially_with_posteriorly_directed_protuberance, 116 setation_on_ventral_metathoracic_femur / absent_or_only_setae_present macrosetae_present 'short,_stout,_cone-shaped_macrosetae_present', 117 cuticular_facets_medially_on_proximal_metathoracic_femur / absent present, 118 ciliate_fringe_ventrally_on_metathoracic_femur_and_tibia / absent present_on_tib_only 'present_on_fem_+_tib', 119 shape_of_metathoracic_tibia / straight arched_laterally arched_medially, 120 length_of_proximal_prothoracic_tarsomere / shorter_or_same_length_as_following_two_tarsomeres_combined longer_than_following_two_tarsomeres_combined, 121 length_of_proximal_mesothoracic_tarsomere / shorter_or_same_length_as_following_two_tarsomeres_combined longer_than_following_two_tarsomeres_combined, 122 length_of_proximal_metathoracic_tarsomere / shorter_or_same_length_as_following_two_tarsomeres_combined longer_than_following_two_tarsomeres_combined, 123 pulvilli / absent present, 124 number_of_dorsal_ridges_on_pulvilli / one two, 125 shape_of_claws / abruptly_angled_distally fairly_straight_throughout, 126 shape_of_tip_of_claws / pointed blunt, 127 development_of_setiform_empodium / 'well-developed_(as_long_as_claws)' reduced_in_length_to_varying_degree minute_or_entirely_absent, 128 anterior_margin_of_wings_in_males / straight C_bulging_anteriorly, 129 upper_calypter / reduced 'well-developed', 130 alula / 'well-developed' reduced_in_size_to_small_lobe 'reduced_entirely,_nearly_straight_wing_margin', 131 cell_r1 / open 'closed,_R1_and_R2+3_meet_apically_and_form_a_stalk_vein_(petiolate)' 'closed,_R1_and_R2+3_meet_directly_at_C', 132 cell_r4 / open closed, 133 cell_r4_shape / R4_and_R5_more_or_less_parallel R4_and_R5_diverging_from_each_other R4_and_R5_approximating_each_other_medially 'R4_and_R5_approximating_each_other_distally_(medially_farthest_apart)' 'R4_and_R5_farthest_apart_in_proximal_one-third', 134 cell_r5 / open closed, 135 cell_m3 / open closed, 136 cell_cup / open closed, 137 cell_d_distal_end / 'base_of_M2_and_m-m_closing_cell_in_nearly_a_single_line' 'only_m-m_closing_cell,_M1_branching_proximally' 'only_base_of_M2_closing_cell,_M3_branching_proximally' 'M2_and_M3_closing_cell,_not_aligned' 'only_base_of_M2_closing_cell,_m-m_absent_(or_at_least_highly_reduced)' 'only_base_of_M2_closing_cell,_distal_M3_absent', 138 'position of r-m in respect to cell d (or cell dm)' / proximal_half distal_half distal_tip, 139 'development of discal medial cell (cell dm)' / absent present, 140 C_extent / 'around_entire_wing_(circumambient)' terminating_at_wing_tip terminating_at_CuA2_and_A1 terminating_at_M2 terminating_at_M1, 141 microtrichia_on_posterior_wing_margin / arranged_in_a_single_plane arranged_in_two_divergent_planes absent, 142 'R2+3 apex' / relatively_straight arching_sharply_anteriorly_in_90¡_angle, 143 R4_apex / terminating_anterior_to_wing_apex terminating_at_wing_apex terminating_posterior_to_wing_apex 'not_reaching_C_(or_wing_margin)', 144 R4_shape / relatively_straight strongly_sinuate, 145 R5_apex / terminating_anterior_to_wing_apex terminating_posterior_to_wing_apex 'not_reaching_C_(or_wing_margin)', 146 'stump vein (supernumerary crossvein) on R4' / absent 'present,_short_stump_vein_not_reaching_R2+3' 'present,_long_stump_vein_reaching_base_of_R2+3', 147 M1_apex / terminating_anterior_to_wing_apex terminating_posterior_to_wing_apex 'not_reaching_C_(or_wing_margin)', 148 M1_and_M2_fusion / separated fused, 149 'M3 development (distal section beyond m-m)' / absent present, 150 'alignment of base of M2 and m-m' / not_aligned aligned_in_a_line_from_anterior_to_posterior 'aligned_in_a_line_from_distal_to_proximal_(nearly_parallel_to_posterior_margin_of_wing)', 151 sculpture_of_tergites / 'smooth,_setae_with_small_sockets_only' 'indented,_setae_with_large_sockets', 152 T1_medially / entirely_sclerotised_medially divided_medially_by_membraneous_area_into_two_halves, 153 T2_length / wider_than_long_or_as_wide_as_long 'longer_than_wide,_less_than_twice_as_long' 'longer_than_wide,_more_than_twice_as_long', 154 T2_shape_dorsally / 'dorsal_surface_smooth,_without_protuberances' dorsal_surface_with_proximal_protuberance, 155 bullae_on_T2 / absent present, 156 'T2-3 development' / tergites_square_to_rectangular 'tergites_forming_a_distinct_''waist''', 157 'T3-6 development' / tergites_entirely_sclerotised tergites_unsclerotised_medially, 158 T6_shape / T6_the_same_as_T5 'T6_cup-shaped,_last_visible_tergite_concealing_remaining_tergites', 159 S2_development / undivided_medially divided_medially_into_two_equal_halves_separated_by_fenestra, 160 'female - lateral macrosetae on T2' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 161 'female - lateral macrosetae on T3' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 162 'female - lateral macrosetae on T4' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 163 'female - lateral macrosetae on T5' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 164 'female - lateral macrosetae on T6' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 165 'female - lateral macrosetae on T7' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 166 'female - segment 7' / normally_developed distinctly_shortened elongated only_S7_reduced_in_size, 167 'female - segments comprising ovipositor' / 8th_and_following 7th_and_following, 168 'female - direction of setae on T8' / posteriorly anteriorly erect, 169 'female - T8 anterior rectangular projection' / 'absent,_no_apodeme_present' 'present,_rectangular_apodeme,_entirely_fused_to_T8' 'present,_rectangular_apodeme,_dissociated_from_T8' 'present,_cylindrical_apodeme,_fused_to_T8', 170 'female - shape of S8' / 'plate-like,_slightly_emarginate_medio-distally' 'plate-like,_with_median_area_elongated' 'proximally_plate-like,_hypogynial_valves_forming_keel_distally' 'keel-like_throughout' 'plate-like,_hypogynial_valves_extending' 'plate-like,_distal_margin_medially_membranous' 'plate-like,_hypogynial_valves_separated_(surrounded_by_membrane)' 'plate-like,_divided_into_two_halves' S8_modified_to_form_part_of_sand_chamber, 171 'female - T9 shape (when distinguishable)' / 'small,_triangular_sclerite' 'wide,_rectangular_sclerite_without_median_protuberance' 'wide,_rectangular_sclerite_with_median_protuberance', 172 'female - T10 shape' / single_sclerite 'tergite_partly_divided_postero-medially' tergite_divided_into_two_heavily_sclerotised_acanthophorites tergite_divided_into_two_heavily_sclerotised_sclerites_which_are_far_apart tergite_divided_into_two_unsclerotised_sclerites T10_lost_or_fused_to_cerci, 173 'female - fusion of T9 and T10' / not_fused partly_fused 'entirely_fused,_sclerites_not_distinguishable', 174 'female - spurs on ovipositor' / absent acanthophorite_spurs_on_T10 spurs_on_cerci, 175 'female - number of spermathecae' / three two one, 176 'female - development of spermathecae' / all_equally_large_spermathecae median_spermatheca_larger_than_lateral_ones median_spermatheca_smaller_than_lateral_ones, 177 'female - length of common spermathecal duct' / 'short,_not_extending_beyond_of_furca' 'long,_extending_beyond_tip_of_furca', 178 'female - length of spermathecal ducts' / short long, 179 'female - ejection apparatus' / absent present, 180 'female - development of spermathecal reservoir' / reservoir_not_differentiated_from_spermathecal_ducts 'reservoir_formed_by_more_or_less_expanded_ducts_to_sac-shaped_reservoir' reservoir_formed_by_more_or_less_expanded_and_coiled_ducts reservoir_spherical, 181 'female - sclerotisation of spermathecal reservoir' / not_sclerotised weakly_sclerotised heavily_sclerotised, 182 'female - length spermathecae' / occupying_at_most_posterior_three_segments occupying_more_than_posterior_three_segments, 183 'female - median sclerite at posterior end of furca' / absent present, 184 'female - development of furca' / a_single_sclerite divided_into_two_lateral_sclerites, 185 'female - furcal apodeme' / absent 'present,_long_plate-like_apodeme' 'present,_short_plate-like_apodeme' 'present,_two_lateral_projections_forming_apodeme', 186 'male - lateral macrosetae on T2' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 187 'male - lateral macrosetae on T3' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 188 'male - lateral macrosetae on T4' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 189 'male - lateral macrosetae on T5' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 190 'male - lateral macrosetae on T6' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 191 'male - lateral macrosetae on T7' / absent 'marginal_(m_l)_setae_present' medial_setae_present, 192 'male - anterior margins of T3-8' / 'straight,_no_apodemes' two_lateral_apodemes, 193 'male - abdominal segments' / 'T1-T8_and_S1-S8' 'T1-T7_and_S1-S7,_T8_+_S8_reduced_to_ring_of_sclerites' 'T1-T7,_T8_entirely_lost' 'T1-T6_and_S1-S6,_T7-T8_and_S7_reduced,_S8_well-developed' 'T1-T8_and_S1-S7,_S8_reduced' 'T1-T7_and_S1-S8,_T8_reduced_to_narrow_sclerite' 'T1-T6_and_S1-S6,_T7-T8_and_S7-S8_reduced_to_ring_of_sclerites', 194 'male - shape of S8' / 'simple,_rectangular_sclerite_or_reduced_to_narrow_sclerite' distal_margin_with_protuberances 'distal_margin_with_flap-like_sclerite', 195 'male - rotation of hypopygium' / not_rotated rotated_by_90¡_to_180¡, 196 'male - direction of hypopygium' / pointing_posteriorly pointing_dorsally, 197 'male - development of epandrium' / 'a_single_sclerite,_fused_more_or_less_entirely_medially' separated_medially_and_joining_proximally separated_medially_and_far_apart_proximally, 198 'male - surstylus on epandrium' / absent 'present,_distal_two-thirds_of_epandrium_form_a_movable_lobe' 'present,_small_epandrial_lobes_connected_to_subepandrial_sclerites', 199 'male - hypandrium shape' / 'well-developed,_rectangular_or_square_sclerite' 'well-developed,_triangular_sclerite' reduced_or_not_differentiated, 200 'male - gonocoxites fusion when hypandrium reduced' / not_fused_medially 'fused_proximo-medially', 201 'male - hypandrium and epandrium' / 'not_approximating,_separated_by_gonocoxites' 'approximating,_but_not_fused_proximally' fused_at_least_partially 'fused,_connected_by_narrow_sclerites_proximally', 202 'male - hypandrium and gonocoxites' / gonocoxites_entirely_free gonocoxites_fused_to_hypandrium 'gonocoxite-hypandrial_complex', 203 'male - epandrium and gonocoxites' / 'gonocoxites_or_gonocoxite-hypandrial_complex_entirely_free' 'gonocoxites_or_gonocoxite-hypandrial_complex_partially_fused_to_epandrium', 204 'male - gonocoxal apodeme' / absent 'present,_short,_at_most_only_slightly_extending_hypopygium_proximally' 'present,_long,_extending_well_into_abdominal_segment_8', 205 'male - development of gonostyli' / absent present, 206 'male - gonostyli position on gonocoxites' / proximal medial distal, 207 'male - lateral processes of gonostyli' / absent present, 208 'male - subepandrial sclerite shape' / no_modifications with_protuberances_medially, 209 'male - subepandrial sclerite distal margin' / 'simple,_straight_margin' distal_projections 'short,_stout_macrosetae', 210 'male - subepandrial sclerite setation' / absent_or_only_a_few_weak_setae_present 'short,_stout_macrosetae', 211 'male - subepandrial sclerite lateral protuberances' / absent present, 212 'male - fusion of cerci' / 'not_fused_(two_distinct_sclerites)' fused_to_a_single_sclerite, 213 'male - lateral ejaculatory process' / absent 'present,_small_triangular_sclerite' 'present,_large_cylindrical_sclerite' 'present,_wide_rectangular_sclerite', 214 'male - functional aedeagus prongs' / one_prong two_prongs three_prongs, 215 'male - dorsal aedeagal sheath length' / 'short,_sperm_sac_entirely_free' 'long,_sperm_sac_entirely_covered', 216 'male - hypandrium posterior margin' / 'simple,_no_particular_projections' long_median_projection, 217 'male - ventral median margin of dorsal aedeagal sheath' / 'weakly_sclerotised,_appearing_incompletely_closed' 'strongly_sclerotised,_appearing_entirely_closed', 218 'male - sperm sac' / appearing_weakly_sclerotised appearing_more_or_less_heavily_sclerotised, 219 'male - lateral ejaculatory process and ventral aedeagal sheath' / 'lateral_ejaculatory_processes_directed_laterally_or_anteriorly,_not_surrounded_by_ventral_aedeagal_sheath' lateral_ejaculatory_processes_directed_posteriorly_and_surrounded_by_ventral_aedeagal_sheath, 220 'male - shape of ejaculatory apodeme' / 'single_plate_of_varying_diameter_and_shape_(two_lateral_surfaces_only)' 'triangular_in_anterior_view_(two_lateral_and_one_dorsal_surface)' 'diamond-shaped_in_anterior_view_(four_lateral_surfaces)' ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220] [ . . . . . . . . . . . . . . . . . . . . . .] Neorhynchocephalus_volaticus 010000--0100030000000003011210100001201000000000000105000000011001001-101200111000101100100000021111100000-0000101000001111100-0000040103-0000000001100000000000000000000010001001032000000000001000002002001200100020100001 Bombylius_major 010000--1001000000000003011010120001100--0000000001100000100011001001-101200011000101100100000011110001000-002010101000111110010000011-0-11000011010000100000000000000003812110001132000000000000010001-01001201000020000100 Poecilognathus_sp._El_Hacha 010000--0000000000000003011010020001100--00300000001011043-0011001001-101200010000001000100010011110000000-002010100000111110010100010-1-110000011100001000000000000000000-0200001111000000000001000001-01001200000020000100 Prorates_sp._Escalante 010000--2111000004000003010010020001101100000000000100001200000000002-112001000000000000101000001100000000-000001100000111110020100010012101000010100001000000000000000000-221????????00000000000000102001021200000010101110 Apsilocephala_longistyla 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0022011020000010000000100110011011200111000001001011001001000201020121011301011011202102111111011210110110-010010201000011110000001020112000100010101000000000011111000002-1220001031000111111100200100-100110001000001011-0 Blepharepium_cajennensis 0012010020010010021000211-11111111101111000001000011021043-00200220121012101001000200002111011010000000112021202020000211111001000001010210010001010100000010000000000001402010000112100200000004010101-10011100000020000101 Brachyrhopala_ruficornis 0021011020110010000001111111111111101111000001000011020043-00200020110011301001000100001111111011100000112011201010200011111000001000000200000001010100000010000000000000402110011112000200000004000000-11011200000030000000 Cabasa_pulchella 0021010020110010000001111011111111101111000001000011120043-00200020111011301001000100001111111010200000112011201010100011111001001000000200000001010100000000000000000000402110011112000300000004000000-11011200000030000000 Chrysopogon_pilosifacies 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0013012020110210000001100111111011100111000000010011001002000102021111011001000000200002111111011200000110-01201010000000011000002001001000000101010100000000000000000020602110001100001200000001010101-11011200010010001001 Lasiopogon_cinctus 001301202011021000000010011111101110011100000001001101100200010202111101100100000020000{23}111111011200000110-01201010000000011000002001001000000101010100000000000000000020602110001100000200000001010101-11011200010010001000 Lissoteles_aquilonius 0011010120110210000000100111111011100111000031010011050002000102020111011011011000200001101111021110010100-002010100000000110000020010010000001010101000000000000000000206021100011010002000000040000020310112002000001010-1 Townsendia_albomacula 0011010020010210000000100111111011100111000001010011010032000102020111010011000000200002111111011100020110-012000200000000110010020010015000000010100000000000000000000206021100011011000000000040000020110112000000001010-1 Damalis_monochaetes 0012011020000010020000211-10111011101111000101000011040023-00100020111010111011000201000100111011100110110-012010211000000110000000010012102000010101000000000000000000000-5200001101100100000001010202012001100000020000000 Damalis_annulata 0013012020000010020000211-10111011101111000101000011040023-00100010111010111011000200000101111011100010110-012010212000000110010000010012102000010101000000000000000000000-5200001121100100000001010002012001200000010000000 Damalis_sp._Palatupana 0010010020000010020001211-10111011101111000101000011040023-00100010121010111011000201001100111011110030110-012010212000101110010000010012102000010101000000000000000000000-5200001121100100000001010002022001200000110001000 Trigonomima_sp._anamaliensis 0020011021010010000000211-10111011101111000101000011011043-00100010111011111011000200000100111011010000110-0120101000000011100100100000{01}5000001010100001000010000000000000-4200001101101200000002-00102022111200000012001000 Holcocephala_abdominalis 0020110020000010020000211-10111011101111000101000011011002000100010111011111011000200000100010021010000110-012010100000000110010010000010000000010101001000000000000000000-4200001101100200000004-00202022111201000012001000 Holcocephala_calva 0020110020000000020000211-10111011101111000101000011011002000100010111011111011000200000100010021010000110-012010100000000110010010000010000000010101000100000000000000000-4200001101100200000004-00202022111201000012001000 Rhipidocephala_sp._Harold_Johnson 0010011021100010000000221-10111011101111000101000011011032000100010111011111011000200000100111021010000110-012010101000011110010010010000000000010101001000010000000000005-0200001101100200000002-00101-11011200000010001000 ; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=ord PolyTcount=MINSTEPS ; TYPESET * UNTITLED = unord: 1-220; END; BEGIN TREES; TRANSLATE 1 Neorhynchocephalus_volaticus, 2 Bombylius_major, 3 Poecilognathus_sp._El_Hacha, 4 Prorates_sp._Escalante, 5 Apsilocephala_longistyla, 6 Hemigephyra_atra, 7 Phycus_frommeri, 8 Nemomydas_brachyrhynchus, 9 Opomydas_townsendi, 10 Pseudonomoneura_hirta, 11 Afroleptomydas_sp._Clanwilliam, 12 Mydas_clavatus, 13 Mitrodetus_dentitarsis, 14 Rhaphiomidas_maehleri, 15 Neorhaphiomidas_sp._Mandurah, 16 Apiocera_aldrichi, 17 Apiocera_painteri, 18 Apoclea_algira, 19 Philodicus_tenuipes, 20 Pogonioefferia_pogonias, 21 Proctacanthus_philadelphicus, 22 Promachus_amastrus, 23 Blepharotes_splendidissimus, 24 Dasophrys_crenulatus, 25 Eichoichemus_pyrrhomystax, 26 Lochmorhynchus_albicans, 27 Megaphorus_pulchrus, 28 Myaptex_brachyptera, 29 Neolophonotus_bimaculatus, 30 Nomomyia_murina, 31 Asilus_crabroniformis, 32 Asilus_sericeus, 33 Satanas_gigas, 34 Lycomya_germainii, 35 Machimus_chrysitis, 36 Machimus_occidentalis, 37 Neomochtherus_pallipes, 38 Afromochtherus_mendax, 39 Dysmachus_trigonus, 40 Philonicus_albiceps, 41 Clephydroneura_sundaica, 42 Clephydroneura_sp._Kepong, 43 Colepia_rufiventris, 44 Congomochtherus_inachus, 45 Heligmonevra_laevis, 46 Neoitamus_cyanurus, 47 Threnia_carbonaria, 48 Tolmerus_atricapillus, 49 Zosteria_rosevillensis, 50 Blepharepium_cajennensis, 51 Brachyrhopala_ruficornis, 52 Cabasa_pulchella, 53 Chrysopogon_pilosifacies, 54 Codula_limbipennis, 55 Cyrtophrys_attenuatus, 56 Deromyia_fuscipennis, 57 Dasypogon_diadema, 58 Austrosaropogon_nigrinus, 59 Lestomyia_fraudiger, 60 Megapoda_labiata, 61 Pseudorus_distendens, 62 Molobratia_teutonus, 63 Neodiogmites_melanogaster, 64 Diogmites_grossus, 65 Pegesimallus_laticornis, 66 Saropogon_dispar, 67 Saropogon_luteus, 68 Senobasis_corsair, 69 Thereutria_amaraca, 70 Cophura_brevicornis, 71 Leptarthrus_brevirostris, 72 Nicocles_politus, 73 Dioctria_atricapillus, 74 Dioctria_hyalipennis, 75 Dioctria_rufipes, 76 Broticosia_paramonovi, 77 Nannodioctria_sp._formicaphaga, 78 Echthodopa_pubera, 79 Dicolonus_simplex, 80 Eudioctria_albius, 81 Andrenosoma_cyrtoxys, 82 Hyperechia_nigripennis, 83 Pilica_formidolosa, 84 Dasyllis_funebris, 85 Proagonistes_praeceps, 86 Atomosia_puella, 87 Aphestia_annulipes, 88 Cerotainia_albipilosa, 89 Goneccalypsis_argenteoviridis, 90 Loewinella_nigripes, 91 Lamyra_gulo, 92 Stiphrolamyra_angularis, 93 Orthogonis_scapularis, 94 Smeryngolaphria_numitor, 95 Laphria_flava, 96 Laphria_aktis, 97 Choerades_bella, 98 Lampria_clavipes, 99 Laxenecera_albicincta, 100 Maira_aenea, 101 Nusa_infumata, 102 Hoplistomerus_nobilis, 103 Laphystia_aegyptiaca, 104 Laphystia_tolandi, 105 Perasis_transvaalensis, 106 Protometer_sp._El_Tuparro, 107 Psilocurus_modestus, 108 Trichardis_effrena, 109 Zabrops_tagax, 110 Leptogaster_cylindrica, 111 Leptogaster_arida, 112 Beameromyia_bifida, 113 Euscelidia_pulchra, 114 Lasiocnemus_lugens, 115 Tipulogaster_glabrata, 116 Afroestricus_chiastoneurus, 117 Cophinopoda_chinensis, 118 Emphysomera_conopsoides, 119 Emphysomera_pallidapex, 120 Michotamia_aurata, 121 Ommatius_incurvatus, 122 Ommatius_tibialis, 123 Acronyches_maya, 124 Bathypogon_nigrinus, 125 Ceraturgus_fasciatus, 126 Myelaphus_melas, 127 Cyrtopogon_lateralis, 128 Cyrtopogon_rattus, 129 Afroholopogon_peregrinus, 130 Dasycyrton_gibbosus, 131 Heteropogon_manicatus, 132 Holopogon_priscus, 133 Metapogon_punctipennis, 134 Nannocyrtopogon_lestomyiformis, 135 Rhabdogaster_pedion, 136 Archilestris_magnificus, 137 Creolestes_nigribarbis, 138 Microstylum_sp._Karkloof, 139 Prolepsis_tristis, 140 Phellus_olgae, 141 Obelophorus_terebratus, 142 Plesiomma_sp._Guanacaste, 143 Stenopogon_sabaudus, 144 Stenopogon_rufibarbis, 145 Acnephalum_cylindricum, 146 Ancylorhynchus_fulvicollis, 147 Ospriocerus_aeacus, 148 Scleropogon_subulatus, 149 Scylaticus_costalis, 150 Sisyrnodytes_sp._Gamka, 151 Tillobroma_punctipennis, 152 Coleomyia_setigera, 153 Hypenetes_stigmatias, 154 Willistonina_bilineata, 155 Daspletis_stenoura, 156 Lycostommyia_albifacies, 157 Gonioscelis_ventralis, 158 Trichoura_sp._Tierberg, 159 Ablautus_coquilletti, 160 Oligopogon_sp._Cape_Recife, 161 Connomyia_varipennis, 162 Stichopogon_elegantulus, 163 Stichopogon_punctum, 164 Stichopogon_trifasciatus, 165 Lasiopogon_aldrichii, 166 Lasiopogon_cinctus, 167 Lissoteles_aquilonius, 168 Townsendia_albomacula, 169 Damalis_monochaetes, 170 Damalis_annulata, 171 Damalis_sp._Palatupana, 172 Trigonomima_sp._anamaliensis, 173 Holcocephala_abdominalis, 174 Holcocephala_calva, 175 Rhipidocephala_sp._Harold_Johnson ; TREE * UNTITLED = [&R] 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